Crocs in JVP: Goniopholis and Metriorhynchus

de Andrade, M. B. and J. J. Hornung. 2011. 'A New Look into the Periorbital Morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and Related Forms' Journal of Vertebrate Paleontology 31(2):352-368 doi: 10.1080/02724634.2011.550353

Abstract
The periorbital morphology of goniopholidids is discussed, exploring the diversity of patterns and the relevance of the data for phylogenetic studies. Revision of material is focused on Goniopholis spp. and aff. Goniopholis spp., from England, Germany, and Belgium, providing a comparative description of their interorbital morphology. Traditional interpretation of the interorbital elements in species of Goniopholis (G. simus, G. baryglyphaeus), where the frontal is interpreted as excluded from the orbit by a prefrontal-postorbital contact in the skull roof, is contested and clarified through the analysis of new specimens, including a morphometric analysis. In Goniopholis, failure to identify the palpebral and its subtle contact with the prefrontal has lead to misinterpretation of elements and structures near the orbit, and the differential preservation/loss of palpebrals explains variability of the orbit in shape and orientation. In all European goniophilidids the frontal reaches the primary orbital border and there is no prefrontal-postorbital contact on the dorsal surface of the skull. Extensive contact of the palpebral with the primary orbital border creates a secondary (functional) orbital border, from which the frontal is excluded in most taxa. The condition is not exclusive of European goniopholidids and is paralleled by protosuchids, peirosaurids, and baurusuchids. At least four main morphological patterns are recognized, revealing a high diversity of European goniopholidids.

Fernandez, M. B., A. P. Carabajal, Z. Gasparini, and G. C. Diaz. 2011. 'A Metriorhynchid Crocodyliform Braincase from Northern Chile' Journal of Vertebrate Paleontology 31(2):369-377 doi: 10.1080/02724634.2011.550361

Abstract
A three-dimensionally preserved metriorhynchid braincase from the Oxfordian of northern Chile is described. The specimen is referred to the metriorhynchid Metriorhynchus cf. M. westermanni. The excellent preservation provides clear sutures and a detailed description, and X-ray computed tomographic (CT) scanning provides internal anatomical details. The general pattern of the orbitotemporal region is consistent with that of the basal thalattosuchian Pelagosaurus typus as described recently. The specimen from northern Chile shares with other metriorhynchids (e.g., Cricosaurus araucanensis, Metriorhynchus westermanni, M. casamiquelai, and Dakosaurus andiniensis) a dorsally exposed laterosphenoid, a laterosphenoid-prootic suture forming a blunt crest separating the supratemporal fenestra into two fossae for muscular attachment, and the quadrate incompletely sutured to the braincase. Thus, these features characterize not only basal but derived Thalattosuchia, as suggested by previous authors. The main difference in the orbitotemporal region is that in the specimen described herein, and in the other metriorhynchids examined, the trigeminal fossa is developed mainly caudal to the trigeminal foramen, whereas in P. typus the fossa is developed rostral and caudal to the trigeminal foramen. CT scanning indicates the presence of enlarged dorsal dural venous sinuses overlying the brain, as it has been described recently in Steneosaurus pictaviensis, and a well-developed sinus within the quadrate. The large foramen ventrolateral to the occipital condyle, which characterizes metriorhynchids, is confirmed as the entry for the internal carotid artery.